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I've never tried it but the idea of constantly holding something just sounds like hell for the carpals.
You don't hold them any more than a horizontal mouse
Wouldn't you thumb it against your palm so that your fingers are always aligned with the buttons, such is the case with laying your hand atop a regular mouse?
Well... I know there's actually three ways to hold a traditional mouse, I saw a video about it once. Which was surprising to me, but mostly because I have big hands and I can only hold a mouse one way.
But for me when I'm holding a horizontal mouse the mouse is "grasped" by my hand as a whole, and I'd say my pinky and thumb are in a way holding the mouse.
With a vertical mouse it's much the same where my whole hand is grasping. I'm not holding it any more with my thumb because the thumb rests on the body of the mouse. In fact as I think about it and type this out I think I have to hold a horizontal mouse more than a vertical, but again it might just be related to hand size.
I use both. Horizontal for gaming so I can have more than two thumb buttons. Vertical for work as it does give my wrist and forearm a "rest."
Edit: realized I didn't necessarily answer your question. For me with the vertical mouse the buttons are just there where they need to be. And I guess clicking is a form of holding...
The G502 has a thumbplatform, so that's where my thumb goes. My pinky can't touch the mouse cause my ring finger is in the way. But its slightly tilted, so its not exactly a horizontal mouse. Not nearly as vertical as this though. I just move it via the top of my palm. A vertical mouse seems like it would be uncomfortable in my right hand but fine with my left (maybe related to mixed handedness, maybe related to the different ways my wrists healed after breaking).